No adapter is configured to be the default adapter. Appletalk was not initialized on any adapter. Services over AppleTalk e.g. Print Server, File Server etc. will not be functional unless a default adapter is specified.
The default adapter setting was deleted by directly accessing the registry instead of by using Network and Dial-up Connections in Control Panel.
Use NCPA to set a default adapter in the Services for Macintosh configuration section by choosing an adapter to be the Default Network.